This course introduces students to a sub-discipline of oceanography that emphasizes the relationships between marine biological and ocean environmental systems. The course will introduce the ecology of planktonic organisms responsible for global ocean carbon cycling, and explain the major dynamical relationships among plankton ecology, population biology and the ocean environment (e.g., critical depth theory, biological pump theory). Applied mathematics is emphasized in these discussions and in the class and lab assignments. In the computer laboratories, students will learn the fundamentals of ocean ecosystem modeling in the R computer language.